There are a lot of things I find interesting about this article, titled “Buddhaguptanatha and the Late Survival of the Siddha Tradition in India,” by David Templeton.

Padmasambhava rupa . . . among the boabs
Padmasambhava rupa . . . among the boabs

Usually, when reading about the history of Indian Buddhism, one finds the narrative ends abruptly in 1193, when the great monastic university N?land? was destroyed by Muslim invaders. Buddhism had been in decline for centuries but, the story goes, the destruction of N?land?, Vikrama??la and the other mahaviharas was the finishing blow. As is often the cause with conventional narratives, this outline is basically true, with caveats.

The end of the great monastic centers does indeed seem to have marked the end of institutionalized, monastic Buddhism in India. But there were, of course, some number of yogins and yoginis living in other settings, and there is much about their fate that we’ll never know. One of the few sources of information we do have is the record of Buddhaguptan?tha, an Indian siddha who showed up in Tibet in the late 16th century and became the primary teacher of the Tibetan Jonang T?ran?tha, who went on to become one of Tibet’s great historians.
Buddhaguptan?tha had belonged to the Natheshvari, a subsect of the Naths, Shaivite Hindu yogins. As Templeton describes, the Natheshvari “combined Buddhist and Hindu teachings . . [they] held Buddhist lineages of instruction within their own Hindu teaching milieu and yet they remained Nath siddhas.” Buddhaguptan?tha reported that he himself developed an entirely Buddhist orientation after having a series of visions of Vajrayogini.
So it seems most likely that Indian Vajrayana Buddhism was absorbed back into the murky, cross-pollinated milieu out of which it arose, and preserved in some form for hundreds of years into the millennium. In fact, in the 1930s one of the Dalai Lama’s teachers, Khunu Lama Tenzin Gyaltsen, supposedly claimed to have met sadhus at Varanasi who, though they outwardly appeared to be Hindus, were secretly Chakrasa?vara practitioners (this according to his student, the translator Christopher Fynn).
One of the more colorful claims of Buddhaguptan?tha’s was that the teachings of Padmasambhava spread to Madagascar. Of course, he seems to have mistaken Jesuits for bhikshus, so he may have been a little confused overall.
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